Gru is the son of Marlena and the Bald Terror (real name Robert), twin brother of Dru, the adoptive father of Margo, Edith, and Agnes, father of Gru Jr., husband of Lucy, and boss of the Minions.
At the beginning of the first film, Gru is an ambitious supervillain who constantly seeks approval from his mother, until the adoption of his daughters convinces him that their happiness is important.
In the second film, Gru leaves his villainous past behind to care for his daughters, but then soon joins forces - unwillingly - with secret agent Lucy Wilde, whom he later marries.
In the third film, after he and Lucy are fired from their jobs at the Anti-Villain League, Gru learns that he has a twin brother, Dru, whom he never met due to their divorced parents raising each child separately. Along with Lucy and the girls, Gru meets Dru at his mansion in Freedonia, and they form a brotherly relationship over the course of the film. Gru and Lucy are also later rehired by the Anti-Villain League.
